Professor Stefan Hendrik Foord (1971–2023)



  • Thinandavha Caswell Munyai School of Life Sciences, University of KwaZulu-Natal, Durban, South Africa





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Peer-reviewed scientific publications

Dippenaar-Schoeman AS, Haddad CR, Lotz LN, Booysen R, Steenkamp RC, Foord SH. 2023. Checklist of the spiders (Araneae) of South Africa. Afr Invertebr. 64(3):221–289.

Azarkina GN, Foord SH. 2023. Further notes on the Afrotropical genus Festucula Simon, 1901 (Araneae, Salticidae). ZooKeys. 1185:295–308.

Ramulifho P, Rivers-Moore N, Foord SH. 2023. Relationships between reference site quality and baetid mayfly assemblages in mountainous streams of the Luvuvhu catchment, South Africa. Water SA. 49(3):251–259.

Lam QD, Rötter RP, Rapholo E, Ayisi K, Nelson WCD, Odhiambo J, Foord SH. 2023. Modelling maize yield impacts of improved water and fertilizer management in southern Africa using Cropping System Model coupled to an Agro–Hydrological Model at field and catchment scale. J Agric Sci. 161(3):356–372.

Bouarakia O, Linden VMG, Joubert E, Weier SM, Grass I, Tscharntke T, Foord SH, Talyor TPJ. 2023. Bats and birds control tortricid pest moths in South African macadamia orchards. Agric Ecosys Environ 352:108527.

Bouarakia O, Anders M, Linden VM, Grass I, Westphal C, Taylor PJ, Foord SH. 2023. Reduced macadamia nut quality is linked to wetter growing seasons but mitigated at higher elevations. J Agric Food Res. 12:100569.

Ferreira NCR, Rötter RP, Bracho-Mujica G, Nelson WC, Lam QD, Recktenwald C, Abdulai I, Odhiambo J, Foord SH. 2023. Drought patterns: their spatiotemporal variability and impacts on maize production in Limpopo province, South Africa. Int J Biometeorol. 67(1):133–148.

Muluvhahothe MM, Joubert E, Foord SH. 2023. Thermal tolerance responses of the two-spotted stink bug, Bathycoelia distincta (Hemiptera: Pentatomidae), vary with life stage and the sex of adults. J Therm Biol. 111:103395.

Dippenaar-Schoeman AS, Foord SH. 2022. Notes on the butterfly Theridiid Episinus marignaci (Lessert, 1933) (Araneae, Theridiidae) from South Africa. Check List. 18(5):1177–1180.

MacFadyen S, Allsopp N, Altwegg R, Archibald S, Botha J, Bradshaw K, Carruthers J, De Klerk H, de Vos A, Distiller G, Foord SH, et al. 2022. Drowning in data, thirsty for information and starved for understanding: A biodiversity information hub for cooperative environmental monitoring in South Africa. Biol Conserv. 274:109736.

Sebata S, Haddad CR, FitzPatrick MJ, Foord SH. 2022. Weak negative responses of spider diversity to short-term ‘kraaling’. Rangeland J. 44(2):61–75.

Kunene C, Foord SH, Scharff N, Pape T, Malumbres-Olarte J, Munyai TC. 2022. Ant Diversity Declines with Increasing Elevation along the Udzungwa Mountains, Tanzania. Diversity (Basel). 14(4):260.

Curveira‐Santos G, Gigliotti L, Silva AP, Sutherland C, Foord SH, Santos‐Reis M, Swanepoel LH. 2022. Broad aggressive interactions among African carnivores suggest intraguild killing is driven by more than competition. Ecology. 103(2):e03600.

Ramulifho PA, Rivers‐Moore NA, Foord SH. 2022. Loss of intermediate flow states only evident when considering sub‐daily flow metrics in a major tributary of the Limpopo basin. Ecohydrology. 15(2):e2381.

Ramulifho PA, Foord SH, Rivers-Moore NA. 2022. Structure and diversity of blackfly assemblages in the Luvuvhu River system, South Africa in response to changing environmental gradients. Hydrobiologia. 849(3):711–723.

Rivers-Moore NA, Ramulifho PA, Foord SH. 2021. Baetid abundances are a rapid indicator of thermal stress and riparian zone intactness. J Therm Biol. 102:103125.

Haddad CR, Foord SH. 2021. Future climate may limit the spread of the Australian house spider Badumna longinqua (Araneae: Desidae) in South Africa. J Arachnol. 49(3):332–339.

Schoeman CS, Foord SH. 2021. Buffer zones maximize invertebrate conservation in a Biosphere Reserve. J Insect Conserv. 25(4):597–609.

Rötter RP, Scheiter S, Hoffmann MP, Pfeiffer M, Nelson WCD, Ayisi K, Taylor P, Feil J-H, Bakhsh SY, Isselstein J, Lindstädter A, Behn K, Westphal C, Odhiambo J, Twine W, Grass I, Merante P, Bracho-Mujica G, Bringhenti T, Lamega S, Abdulai I, Lam QD, Anders M, Linden V, Weier S, Foord SH, Erasmus B. 2021. Modeling the multi‐functionality of African savanna landscapes under global change. Land Degrad Develop 32:2077–2081.

Haddad CH, Foord SH, Whitehead L. 2021. Tussock circumference, land use type and drought variably influence spider assemblages associated with Hyparrhenia hirta grass tussocks. Afr Entomol. 29(1):150–164.

Munyai TC, Khoza LR, Foord SH. 2021. Hidden just beneath our feet: subterranean ants of the KwaZulu-Natal midlands. Afr J Ecol. 59(2):554–560.

Muluvhahothe MM, Joseph GS, Seymour CL, Munyai TC, Foord SH. 2021. Repeated surveying over 6 years reveals that fine-scale habitat variables are key to tropical mountain ant assemblage composition and functional diversity. Sci Rep. 11:56.

Dippenaar-Schoeman AS, Foord SH. 2020. Revision of the Afrotropical crab-spider genus Parabomis Kulczyński, 1901 (Araneae: thomisidae). Zootaxa. 4899(1):161–174.

Shivambu CT, Shivambu N, Lyle R, Jacobs-Venter R, Kumschick S, Foord SH, Robertson MP. 2020. Tarantulas (Araneae: Theraphosidae) in the pet trade in South Africa. Afr Zool. 55(4):323–336.

Schoeman CS, Toussaint DC, Tshililo P, Foord SH, Hamer M. 2020. Checklist of the pimeliine darkling beetles of the Vhembe Biosphere Reserve, South Africa (Coleoptera: Tenebrionidae: Pimeliinae). Bothalia. 50(1).

Foord SH, Dippenaar-Schoeman AS, Haddad CR, Lyle R, Lotz L, Sethusa T, Raimondo D. 2020. The South African National Red List of spiders: patterns, threats, and conservation. J Arachnol. 48(2):110–118.

Ramulifho PA, Foord SH, Rivers-Moore NA. 2020. The role of hydro-environmental factors in Mayfly (Ephemeroptera, Insecta) community structure: identifying threshold responses. Ecol Evol. 10(14):6919–6928.

Linden B, Foord SH, Horta-Lacueva QJ, Taylor PJ. 2020. Bridging the gap: how to design canopy bridges for arboreal guenons to mitigate road collisions. Biol Conserv. 246:108560.

Dippenaar-Schoeman AS, Wiese L, Foord SH, Haddad CR. 2020. A list of spider species found in the Addo Elephant National Park, Eastern Cape province, South Africa. Koedoe. 62(1):a1578.

Cuthbert RN, Sithagu R, Weyl OL, Wasserman RJ, Dick JT, Callaghan A, Froneman PW, Foord SH, Dalu T. 2020. Water volume differentially modifies copepod predatory strengths on two prey types. Limnologica. 81:125747.

Schoeman CS, Hahn N, Hamer M, Foord SH. 2020. Regional invertebrate cross-and within-taxon surrogacy are scale and taxon dependent. Trans R Soc S Afr. 75:23–32.

Dippenaar-Schoeman AS, Foord SH. 2019. New records of Cladomelea from South Africa, including the first records of C. longipes (O. Pickard-Cambridge, 1877) (Araneae, Araneidae) outside its type locality. Check List. 15(5):1071–1075.

Foord SH, Dippenaar-Schoeman AS, Haddad CR, Schoeman C, Hahn N, Lyle R. 2019. Spider checklist for the Blouberg, in the Vhembe Biosphere Reserve, South Africa. Bothalia. 49(1):a2455.

Dean WRJ, Seymour CL, Joseph GS, Foord SH. 2019. A review of the impacts of roads on wildlife in semi-arid regions. Diversity 11:81.

Cardoso P, Shirey V, Seppälä S, Henriques S, Draney ML, Foord SH, Gibbons ATL, Gomez A, Kariko S, Malumbres-Olarte J, et al. 2019. Globally distributed occurrences utilised in 200 spider species conservation profiles (Arachnida, Araneae). Biodivers Data J. 7:e33264.

Bishop TR, Parr CL, Gibb H, van Rensburg BJ, Braschler B, Chown SL, Foord SH, Lamy K, Munyai TC, Okey I, et al. 2019. Thermoregulatory traits combine with range shifts to alter the future of montane ant assemblages. Glob Change Biol. 25(6):2162–2173.

Joseph GS, Muluvhahothe MM, Seymour CL, Munyai TC, Bishop TR, Foord SH. 2019. Stability of Afromontane ant diversity decreases across an elevation gradient. Glob Ecol Conserv. 17:e00596.

Haddad CR, de Jager LJC, Foord SH. 2019. Habitats and cardinal directions are key variables structuring spider leaf litter assemblages under Searsia lancea. Pedobiologia (Jena). 73:10–19.

Seppälä S, Henriques S, Draney ML, Foord SH, Gibbons AT, Gomez LA, Kariko S, Malumbres-Olarte J, Milne M, Vink CJ, et al. 2018. Species conservation profiles of a random sample of world spiders IV: Scytodidae to Zoropsidae. Biodivers Data J. 6:e30842.

Seppälä S, Henriques S, Draney S, Foord S, Gibbons A, Gomez L, Kariko S, Malumbres-Olarte J, Milne M, Vink C, et al. 2018. Species conservation profiles of a random sample of world spiders III: Oecobiidae to Salticidae. Biodivers Data J. 6:e27004.

Seppälä S, Henriques S, Draney M, Foord SH, Gibbons A, Gomez L, Kariko S, Malumbres-Olarte J, Milne M, Vink C, et al. 2018. Species conservation profiles of a random sample of world spiders II: Gnaphosidae to Nemesiidae. Biodivers Data J. 6:e26203.

Dippenaar-Schoeman AS, Haddad CR, Lyle R, Lotz LN, Foord SH, Jocque R, Webb P. 2018. South African National Survey of Arachnida: A checklist of the spiders (Arachnida, Araneae) of the Tswalu Kalahari Reserve in the Northern Cape province, South Africa. Koedoe. 60(1):a1486.

Foord SH, Swanepoel LH, Evans SW, Schoeman CS, Erasmus BFN, Schoeman MC, Keith M, Smith A, Mauda EV, Maree N, et al. 2018. Animal taxa contrast in their scale-dependent responses to land use change in rural Africa. PLoS One. 13(5):e0194336.

Seppälä S, Henriques S, Draney M, Foord SH, Gibbons A, Gomez L, Kariko S, Malumbres-Olarte J, Milne M, Vink C, et al. 2018. Species conservation profiles of a random sample of world spiders I: Sgelenidae to Filistatidae. Biodivers Data J. 6:e23555.

Grass I, Meyer S, Taylor PJ, Foord SH, Hajek P, Tscharntke T. 2018. Pollination limitation despite managed honeybees in South African macadamia orchards. Agric Ecosyst Environ. 260:11–18.

Joseph GS, Seymour CL, Coetzee BWT, Ndlovu M, Deng L, Fowler K, Hagan J, Brooks BJ, Seminara JA, Foord SH. 2018. Elephants, termites and mound thermoregulation in a progressively warmer world. Landsc Ecol. 33(5):731–742.

Mauda EV, Joseph GS, Seymour CL, Munyai TC, Foord SH. 2018. Changes in landuse alter ant diversity, assemblage and dominant functional groups in African savannas. Biodivers Conserv. 27(4):947–965.

Joseph GS, Mauda EV, Seymour CL, Munyai TC, Dippenaar-Schoeman AS, Foord SH. 2018. Land use change in savannas disproportionally reduces functional diversity of invertebrate predators at the highest trophic levels: spiders as an example. Ecosystems (NY). 21(5):930–942.

Ramulifho PA, Rivers-Moore NA, Dallas HF, Foord SH. 2018. A conceptual framework towards more holistic freshwater conservation planning through incorporation of stream connectivity and thermal vulnerability. J Hydrol (Amst). 556:173–181.

Simba LD, Foord SH, Thébault E, van Veen F, Joseph GS, Seymour CL. 2018. Indirect interactions between crops and natural vegetation through flower visitors: The importance of temporal as well as spatial spillover. Agric Ecosyst Environ. 253:148–156.

Modiba RV, Joseph GS, Seymour CL, Fouché PSO, Foord SH. 2017. Restoration of riparian systems through clearing of invasive plant species improves functional diversity of Odonate assemblages. Biol Conserv. 214:46–54.

Joseph GS, Seymour CL, Foord SH. 2017. The effect of infrastructure on the invasion of a generalist predator: pied crows in southern Africa as a case-study. Biol Conserv. 205:11–15.

Joseph GS, Seymour CL, Coetzee BWT, Ndlovu M, De La Torre A, Suttle R, Hicks N, Oxley S, Foord SH. 2016. Microclimates mitigate against hot temperatures in dryland ecosystems: termite mounds as an example. Ecosphere. 7(11):e01509.

Foord SH, Dippenaar-Schoeman AS, Jocqué R, Haddad CR, Lyle R, Webb P. 2016. South African National Survey of Arachnida: A checklist of the spiders (Arachnida, Araneae) of the Lekgalameetse Nature Reserve, Limpopo Province, South Africa. Koedoe. 58(1):a1405.

Bishop TR, Robertson MP, Gibb H, van Rensburg BJ, Braschler B, Chown SL, Foord SH, Munyai TC, Okey I, Tshivhandekano PG, et al. 2016. Ant assemblages have darker and larger members in cold environments. Glob Ecol Biogeogr. 25(12):1489–1499.

Traoré AN, Mulaudzi K, Chari GJ, Foord SH, Mudau LS, Barnard TG, Potgieter N. 2016. The impact of human activities on microbial quality of rivers in the Vhembe District, South Africa. Int J Environ Res Public Health. 13(8):817.

Foord SH, Dippenaar‐Schoeman AS. 2016. The effect of elevation and time on mountain spider diversity: a view of two aspects in the Cederberg mountains of South Africa. J Biogeogr. 43(12):2354–2365.

Foord SH, Fouche PSO. 2016. The response of instream animal communities to a short term extreme event and longer term cumulative impacts in a strategic water resource area, South Africa. Afr J Aquat Sci. 41(1):29–40.

Haddad CH, Foord SH, Fourie R, Dippenaar-Schoeman AS. 2015. Effects of a fast-burning spring fire on the ground-dwelling spider assemblages (Arachnida: Araneae) in a central South African grassland habitat. Afr Zool. 50(4):281–292.

Dippenaar-Schoeman AS, Haddad CH, Foord SH, Lyle R, Lotz LN, Marais P. 2015. South African National Survey of Arachnida (SANSA): review of current knowledge, constraints and future needs for documenting spider diversity (Arachnida: Araneae). Trans R Soc S Afr. 70(3):245–275.

Azarkina GN, Foord SH. 2015. A review of three Tusitala (Araneae: Salticidae) species from southern Africa, with a new synonymy and description of a new species from Botswana. Afr Invertebr. 56(2):258–307.

Munyai TC, Foord SH. 2015. Temporal patterns of ant diversity across a mountain with climatically contrasting aspects in the tropics of Africa. PLoS One. 10(3):e0122035.

Munyai TC, Foord SH. 2015. An inventory of epigeal ants of the western Soutpansberg mountain range, South Africa. Koedoe. 57(1):1244.

Foord SH, Gelebe V, Prendini L. 2015. Effects of aspect and altitude on scorpion diversity along an environmental gradient in the Soutpansberg, South Africa. J Arid Environ. 113:114–120.

Azarkina GN. Foord SH. 2014. A revision of the Afrotropical species of Festucula Simon, 1901 (Araneae: Salticidae). Afr Invertebr. 55(2):351–375.

Azarkina GN, Foord SH. 2013. Redescriptions of poorly known species of jumping spiders (Araneae: Salticidae) from South African and Namibia. Zootaxa. 3686:165–182.

Foord SH, Dippenaar-Schoeman AS, Stam EM. 2013. Surrogates of spider diversity, leveraging the conservation of a poorly known group in the Savanna Biome of South Africa. Biol Conserv. 161: 203–212.

Haddad CR, Dippenaar-Schoeman AS, Foord SH, Lotz LN, Lyle R. 2013. The faunistic diversity of spiders (Arachnida: Araneae) of the South African Grassland Biome. Trans R Soc S Afr. 68:97–122.

Dippenaar-Schoeman A, Van den Berg A, Lyle R, Haddad C, Foord S, Lotz L. 2013. The diversity of South African spiders (Arachnida: Araneae: Documenting a National Survey). Suid-Afrikaanse Tydskrif vir Natuurwetenskap en Tegnologie. 19;32(1):375.

Munyai TC, Foord SH. 2012. Ants on a mountain: spatial, environmental and habitat associations along an altitudinal transect in a centre of endemism. J Insect Conserv. 16(5):677–695.

Schoeman CS, Foord SH. 2012. Deciding on an appropriate scale for conservation activities: partitioning alpha and beta ant diversity in North-West province, South Africa (Formicidae: Hymenoptera). Trans R Soc S Afr. 67(1):1–10.

Schoeman CS, Foord SH. 2012. A checklist of epigaeic ants (Hymenoptera: Formicidae) from Marakele National Park, Limpopo, South Africa. Koedoe 54(1):a1030.

Foord SH, Dippenaar-Schoeman AS, Haddad CR, Lotz LN, Lyle R. 2011. The faunistic diversity of spiders (Arachnida, Araneae) of the Savanna Biome in South Africa. Trans R Soc S Afr. 66(3):170–201.

Javed SMM, Foord SH, Tampal F. 2010. A new species of Hersilia Audouin, 1826 (Araneae: Hersiliidae) from India, with notes on its natural history. Zootaxa. 2613(1):40–50.

Muelelwa MI, Foord SH, Dippenaar-Schoeman AS, Stam EM. 2010. Towards a standardized and optimized protocol for rapid biodiversity assessments: spider species richness and assemblage composition in two savanna vegetation types. Afr Zool. 45(2):273–290.

Foord SH, Mafadza M, Dippenaar-Schoeman AS, Van Rensburg BJ. 2008. Micro-scale heterogeneity of spiders (Arachnida: Araneae) in the Soutpansberg, South Africa: a comparative survey and inventory in representative habitats. Afr Zool. 43(2):156–174.

Foord SH. 2008. Cladistic analysis of the family Hersiliidae (Arachnida, Araneae) of the Afrotropical region with the first records of Murricia, and the description of a new genus, Prima, from Madagascar. J Afr Zool. 4:111–141.

Foord SH, Dippenaar-Schoeman AS. 2006. A revision of the Afrotropical species of Hersilia audouin (Araneae: hersiliidae). Zootaxa. 1347(1):1–92.

Dippenaar-Schoeman AS, Van den Berg AM, Van den Berg MA, Foord SH. 2005. Spiders in avocado orchards in the Mpumalanga Lowveld of South Africa: species diversity and abundance (Arachnida: Araneae). Afr Plant Prot. 11:8–16.

Foord SH, Dippenaar-Schoeman AS. 2005. First records of the genus Neotama Baehr & Baehr (Araneae: Hersiliidae) from the Afrotropical Region. Afr Invertebr. 45:125–132.

Foord SH, Dippenaar-Schoeman AS. 2005. A revision of the Afrotropical species of Hersiliola thorell and Tama simon with description of a new genus Tyrotama (Araneae: hersiliidae). Afr Entomol. 13:255–279.

Foord SH, Ferguson JWH, Van Jaarsveld AS. 2003. Coleopteran assemblages in Afromontane grasslands reflect fine-scale variation in vegetation. Environ Entomol. 32(4):797–806.

Ferguson JWH, Van Jaarsveld AS, Johnson R, Bredenkamp GJ, Foord SH, Britz M. 2003. Rodent-induced damage to pine plantations: a South African case study. Agric Ecosyst Environ. 95(1):379–386.

Foord SH, Dippenaar-Schoeman AS, Van der Merwe M. 2002. A check list of the spider fauna of the Western Soutpansberg, South Africa (Arachnida: araneae). Koedoe. 45(2):35–43.

Foord SH, Ferguson JWH, Van Jaarsveld AS. 2002. Endemicity of Afromontane grasshopper assemblages and its implications for grassland conservation. Afr J Ecol. 40(4):318–327.

Bredenkamp GJ, Ferguson JWH, Foord SH, De Frey WH. 1999. Floristic assessment of an Afromontane grassland fragmentation experiment in a pine plantation. Afr J Ecol. 37(1):49–60.

Foord SH, Van Aarde RJ, Ferreira SM. 1994. Seed dispersal by vervet monkeys in rehabilitating coastal dune forests at Richards Bay. S Afr J Wildl Res. 24:56–59.

Edited books and book chapters

Foord SH, Dippenaar-Schoeman AS, Haddad CH. 2011. South African Spider Diversity: African Perspectives on the Conservation of a Mega-diverse Group. In: Grillo O, Venora G (eds.), Changing Diversity in Changing Environment. Rijeka: Intech. pp.163–184.

Dippenaar-Schoeman AS, Foord SH, Haddad CH. 2013. Spiders of the Savanna Biome. Thonoyandou: University of Venda, Pretoria: Agricultural Research Council.




How to Cite

Munyai TC. Professor Stefan Hendrik Foord (1971–2023): Obituary. Afr. Entomol. [Internet]. 2024 Feb. 29 [cited 2024 Jul. 24];32. Available from: